Can anyone give me an explanation of how they work? I have the combo green boxes appearing around my cards in my hand in battle, but I always end up using one skill or the other, not actually combining the glyphs for a combo.
You can tap the "R" key to make combos. Tap "R" and then the two corresponding hotbar slot numbers to combine those two glyphs into a combo. Alternately, use the mouse to drag and drop glyphs on top of each other to make a combo.